颚化符`~`在枚举中使用

Iv'e看看这个 answer

似乎无法理解这行代码:

CultureInfo[] cinfo = CultureInfo.GetCultures(CultureTypes.AllCultures & ~CultureTypes.NeutralCultures);

public enum CultureTypes
{
    NeutralCultures = 1,
    SpecificCultures = 2,
    InstalledWin32Cultures = 4,
    AllCultures = 7,
    UserCustomCulture = 8,
    ReplacementCultures = 16,
    [Obsolete("This value has been deprecated.  Please use other values in CultureTypes.")]
    WindowsOnlyCultures = 32,
    [Obsolete("This value has been deprecated.  Please use other values in CultureTypes.")]
    FrameworkCultures = 64,
}

CultureInfo[] cinfo = CultureInfo.GetCultures(CultureTypes.AllCultures | 
CultureTypes.NeutralCultures | 
CultureTypes.SpecificCultures | 
CultureTypes.InstalledWin32Cultures | 
CultureTypes.UserCustomCulture | 
CultureTypes.ReplacementCultures );